| Authorship |
Williams et al., 2025:
584, figs. 62-70
|
| Forms |
w.q.m. |
| Bioregion |
Neotropic |
| Locality |
COSTA RICA |
| Etymology |
The species epithet is derived from Frank Herbert’s science fiction novel Dune. In the novel, “Usul” is the secret name given to the protagonist, Paul Atreides, by the Fremen, the desert dwellers of the planet Arrakis. This name references the species’ moderate to intense blue iridescence, alluding to the “blue-within-blue” eyes that Paul acquires after joining the Fremen and consuming the spice melange. The term “Usul” translates to “the strength of the base of the pillar” and is derived from the Arabic word usul, the plural form of asl, which means “origin,” “root,” or “principle.” |
| Primary type information |
Primary type material: holotype worker. Primary type locality: Costa Rica: San José, 11km SSE Santa María, 9.55301 -83.94523 ±10m, elev. 1950m, 24.vi.2015, cloud forest, sifted litter, ADMAC#Wa-E-04-1-49. Primary type depository: USNM. Primary type specimen: USNMENT01131798. |
| Secondary type information |
Secondary type material: 5 paratype workers, 1 paratype queen, 1 paratype male. Secondary type locality: same as for holotype. Secondary type depositories: MNCR, USNM. Secondary type specimens: USNMENT01131109, USNMENT01131799, USNMENT01131800, USNMENT01131802, USNMENT01131804, USNMENT01131819, USNMENT01131831. |
