Primary type information:
Primary type material: holotype male. Primary type locality: Germany: Rhineland Palatinate, Enspel Oilshale; Enspel Formation, Upper Oligocene, Enspel Formation, Upper Oligocene, MP 28; (24.56–24.79 Ma, Mertz et al. 2007), layer S16. Primary type depository:
NHMM. Primary type specimen: NHMM-PE1997/5513-LS.