Primary type information:
Primary type material: holotype alate queen. Primary type locality: Germany: Rhineland-Palatinate, Enspel Oilshale, Enspel Formation, Upper Oligocene, MP 28; (24.56–24.79 Ma,
Mertz et al. 2007), layer S16. Primary type depository:
NHMM. Primary type specimen: NHMM-PE2009/5826-LS.