Primary type information:
Primary type material: holotype queen. Primary type locality: U.S.A.: Montana, Canyon Ferry Reservoir; Lower Oligocene (Rupelian, ca. 32.0 ± 0.1 Ma). Primary type depository:
MOR. Primary type specimen: MOR-IV-1465.
Secondary type information:
Secondary type material: 3 paratype queens. Secondary type locality: same as for holotype. Secondary type depository:
MOR. Secondary type specimens: MOR-IV-1047, MOR-IV-1107, MOR-IV-1156.
Type notes:
The age range of 32 ± 0.1 Ma is probably best regarded as the uncertainty of an upper bound, not as the total deposit age range. This date is derived from uranium-lead dating of a tuff located 188 meters below the insect horizon, and as such the fossiliferous layer may be considerably younger, for example of Arikareean age (29.5-18.5 Ma) based on possible association with nearby vertebrate fossils (CoBabe, Chamberlain, Ivie & Giersch 2002. Rocky Mountain Geology 37(1): 13-30).